A great link, but I will say that it's not complete as of yet. I know a few people still working on the arthropod side of the tree.
Yes, I saw that Neoptera is just a hideously large polytomy, for instance, and Phthiraptera is just a list... but nevertheless, as randman is going to show us groupings in the general pattern, I think this will still be a sufficiently resolved source, as long as we keep in mind that it is a consensus tree.
Mostly because it's an overly generalized statement which is unsupportable. The thing is, genetic isolation says NOTHING about diversity. An isolated population can have more, less, or an equal amount of variation, because there's no causal connection between the concepts.
I am not convinced that his argument has any bearing on the strength of any given phylogenetic tree as evidence for evolutionary theory even if there was such a link, but at the moment it's getting to late for me to think about it properly.
I have already substantiated it. You just don't seem capable of grasping the basic concepts involved. Sequential speciation is how evos imagine the higher taxa involved.
I don't understand why you keep mentioning this, as that is nothing I am disputing. What I am disputing is your claim that you have shown groupings in any phylogenetic tree -- even the global consensus tree to which I linked -- which would not be predicted by evolutionary theory. If you have shows any such groupings, please link to them again, as I must have missed them.
This involves subgroup speciation, which decreases genetic variability. Genetic variability is considered by evos themselves to be critical to an organisms ability to survive and evolve further.
I agree that this is generally the case, though need not be the case if subgroup isolation is the result of allopolyploidy, for instance.
However, even an isolated subgroup which has a reduced genetic variation compared to the mother population will be subject to random point mutations and other mechanisms that will serve to increase variation (supposing no extinction occurs) over time. This is called a bottle neck, and is known to have happened in a great many still extant taxa. Why, at least two species in the group of birds I work on (Calidrinae) are known to have gone through bottle necks in recent time (the Dunlin and the Red Knot), but then managed to diverge again. Remarkable, they constitute two of the three Calidrine shorebirds that show sufficient morphological variation even today to be subdivided into subspecies (the third being the Broad-billed Sandpiper.
There are other reasons. Macroevolution, if it occurred, seems to be an irreversible process buy allelic mutation being random has no direction.
This statement contains no contradiction. Certainly the same position in a sequence can and does mutate multiple times within a lineage, especially in sections of the DNA that is less conservative. This is a well-known fact in evolutionary theory, and has been for at least as long as we have used genetic sequences for phylogenetic analysis. However, the chance of every single mutation in a lineage being reversed at the same time and thereby producing the exact same sequence as in a removed ancestral organism increases (linearly?) with the temporal distance between the two taxa, and (unpredictably?) with the effect the mutations of the ancestral sequence have on the biology of the organism in question. Therefore, when enough change has occurred in a lineage so that we would classify it as a different taxa from its ancestor, the chances of all these mutations being reversed at the same time in the same descendant further on is incredibly small, but certainly within the realms of possibility.
This, incidentally, is the reason why different genes are used for resolution on different taxonomic levels when we do phylogenetics. Genes known to have a high rate of mutation (
i.e., genes that evolve fast) will give resolution only at low taxonomic levels, whereas more conservative ones will give resolution only at higher taxonomic levels. For instance, if I were to construct a phylogeny of the whole of Neoptera and was forced to use only one gene, I would not use COI as that has too high a mutation rate, and would result in a large polytomy. While this is, of course, also data, it is not data that can be used in any way, and thus I would chose another gene.
Macroevolution does not appear to be happening any more.
Would you care to support this assertion?
So how do you explain convergent evolution since the probability is so low for mutations to repeat?
This statement does not contain a contradiction. Convergent evolution does not imply that any given gene in two distantly related lineages have mutated into the same gene. It implies that two distantly related lineages show great morphological similarities, but these may have come about by vastly different genetic routes. For instance, if two groups of birds have both lost their colour and become all white, this could be because one lineage has had a mutation in the genes that produce pigment, whereas the other has one in the gene that transports the pigments from their point of manufacture to the feathers. The two would be said to have undergone convergent evolution, but their genetic sequences would still be very different.
Take a look a map of phylogenetic trees carefully. First note the common ancestor is always missing. Second, macroevolution appears to have stopped. No new kingdoms and phyla for a long time. No duplicates of a bacteria evolving again from a different source. No reversibility. Macroevolution, and this assumes common, descent appears to happen in irreversible pulses that peter out and end.
While I appreciate that you have condensed your confusion, allowing me to address all of it without having to use the quote button multiple times, I'd rather you'd forgo this service and instead expound on your novel theories on how evolution, phylogeny and taxonomy works. If I may make a suggestion, I would recommend that you answer specific points I make, rather than just sum up everything at the end. This gives at least the impression that you have read anything I write.
Now:
Take a look a map of phylogenetic trees carefully. First note the common ancestor is always missing.
No, it isn't. The common ancestor is the node where two lineages converge. This is always present, or you wouldn't have a tree. You would have a bunch of parallel lines.
Further, the use of outgroups constrains the ingroup so that the common ancestor of all studied taxa is included in the analysis. This is evident in all published and unpublished trees I have ever seen.
In morphological trees and genetic trees that include character tracking, the specifics of any given common ancestor between two lineages is clearly denoted in the tree or, at least, in the text. In some morphological trees, these internal nodes even have labels attached to them that suggest known taxa that show these characteristics and may therefore be a candidate for the direct ancestor of those lineages.
Feel free to show me a phylogenetic tree that doesn't include common ancestors. As I said, I have access to most of the major journals where phylogenies would be published, so even a reference would be enough. And if I can't get it, I am sure I could get help from someone else in the thread, or simply write the author and ask for it. So don't feel constrained on my behalf when selecting examples.
Second, macroevolution appears to have stopped. No new kingdoms and phyla for a long time.
As I have pointed out to you before, and as Dinwar so brilliantly explained in another thread, this is because these taxa do not exist in nature. There is no such thing as a "phylum". These are merely labels we attach to groupings of organisms which show certain common characteristics. The longer this group has been around, the more likely it is to have a label at a higher exclusive rank. The lack of recent new phyla is because this label is one that is attached only to groups that have a long evolutionary and geological history as an entity separate from all other phyla. Nothing that is today classified into a phylum will ever involve into something that would be classified into another phylum, because that is how taxonomy works. All Chordata will always be Chordata, even if they evolve to lose their spines.
No duplicates of a bacteria evolving again from a different source.
You mean that no organism has ever evolved into a bacterium? Yes, that is certainly true, but it is a tautology, as a given taxonomic category ideally contains only the descendants of a single common ancestor. Even if your descendants evolve into something that for all practical purposes is indistinguishable from a bacteria, you would still not be classified as one, as you do not have a shared exclusive evolutionary history with other bacteria.
There are countless examples of reversibility, if by this you mean that a certain feature is first evolved, and then lost again in subsequent generations. If you imply a different meaning, please explain.
Macroevolution, and this assumes common, descent appears to happen in irreversible pulses that peter out and end.
Again, this is easy for a beginner to believe because any phylogenetic tree can give the evolutionary history of the included taxa only up until the date the oldest included individual was collected. As soon as a given individual is dead, it will no longer have further descendants, and evolution along that individual's lineage will stop there. Naturally, if an included individual had descendants before the collection date, or DNA was collected non-lethally, that individual would still have descendants, but their future evolution will not be picked up by the DNA as it is, by definition, in the future.
You guys act like you never even heard the NeoDarwinian, etc,....
Again, you are too caught up in what others know and don't know. For the purposes of this discussion, I am entirely uninterested in knowing what Sceptic Tank, Dinwar, or anyone else arguing against you knows or doesn't know about evolution. I am interested only in what
you know about it. I am a professional biologist, and work with evolutionary theory every day. You claim to be able to show that what I am doing is either fraudulent or at least pointless. I do not care to be neither a fraud, nor a waste of taxpayers' money, so if you truly have this data, you would genuinely do me a great service if you presented it, and thus allowed me to review my career choices and perhaps change tracks into something more useful. I have, for instance, recently taken up pottery, which I find very rewarding.
Why you continue to be reluctant to do me this service is beyond me. Had I known you personally, I might even have claimed it to be rude.
Learn what your side believes first; then take a little while to learn what others believe, and then we can have an intelligent discussion.
I will agree that under the circumstances, it is hard to have an intelligent discussion here.